bees in the hive entrance two bees in the hive entrance bee collected pollen Yael and Tal do a homing experiment

Lab Members


Prof. Sharoni Shafir

Sharoni has been the director of the B. Triwaks Bee Research Center since 1998. He advocates a multidisciplinary approach to promoting bee health and bee pollination services. He is especially interested in bee cognition, specifically aspects of perception and decision making that relate to foraging and pollination. He has a BA and PhD from Stanford University, an MS from University of Florida, and was a postdoctoral fellow at The Ohio State University.

Dr. Harmen P. Hendriksma

Postdoctoral Fellow

Harmen is a devoted beekeeper and researcher. He studied ecosystem biology at the University of Wageningen, in Holand. Thereafter, he was a professional beekeeper for 4 years until he started his PhD in Bayreuth and Wuerzburg in Germany. His PhD research was on honey bee risk assessment with transgenic maize. Harmen started his postdoc position in Rehovot, Israel in August of 2012 on the topic of the nutritional balancing done by honey bees.

  • Harmen's publications
  • Read more about Harmen's PhD research.
  • harmenhendriksma@gmail.com

    Haim Kalev

    Lab Manager and Beekeeper

    Haim has a B.Sc. in Agriculture from the Hebrew University of Jerusalem. He has been a technician and beekeeper at the B. Triwaks Bee Research Center since 1997. Haim is very involved in education and collaboration with the international community of beekeepers. He has led many courses in subjects such as beekeeping for beginners, modern apiculture management (honey by-products and pollination), commercial beekeeping, and artificial insemination of queens. His courses take place all over the world, including Eritrea, Nigeria, Ethiopia, Uzbekistan, Rwanda, and El Salvador.

    kalev@agri.huji.ac.il

    Amir Raz

    PhD Student

    Co-supervisor: Prof. Martin Goldway and Dr. Raffi Stern

    Thesis: Pollination in apricots (Prunus armeniaca): honey bee foraging behavior and genetic aspects of the fertilisation mechanism

    amirr@migal.org.il

    Yael Garbian

    Ph.D Student

    Co-supervisor: Prof. Ilan Sela

    Yael Garbian has a B.Sc. in Agriculture and Biotechnology from the Hebrew University. She did her M.Sc. in Biology at Ben Gurion University. Her M.Sc. research focused on complex 1 of oxidative phosphorylation in mitochondria. She is currently doing her Ph.D research, at the Hebrew University's Faculty of Agriculture, on the molecular mechanisms of honey bee diseases.

    garbiland@gmail.com

    Reut Nyska

    M.Sc. Student

    Co-supervisor: Dr. Doron Schneider

    Thesis: The gametophytic self-incompatibility system in loquat (Eriobotrya japonica) and the foraging behavior of honey bees

    reutnyska@gmail.com

    Yael Katz

    M.Sc. Student

    Co-supervisor: Dr. Arnon Dag

    Yael has a B.Sc. in Agriculture from the Hebrew University of Jerusalem. Her M.Sc. research focuses on the importance of fatty acids in honey bee nutrition, especially on learning and memory. Yael uses methods such as proboscis extension response (PER).

    yaelkat@gmail.com

    Tal Erez

    M.Sc. Student

    Co-supervisor: Dr. Oz Barazani

    Tal has a B.Sc. in Agriculture from the Hebrew University of Jerusalem. Her M.Sc. research focuses on preferences of honey bees and decisions that they make when they are pollinating two morphs of the same flower.

    tal.erez2@mail.huji.ac.il

    Tania Masci

    Laboratory Technician

    Tania has worked at the Faculty of Agriculture since 1997. She is a professional bee brain surgeon! She is currently working with Yael Katz on the effect of fatty acid nutrition on learning and memory. She has a B.Sc. in Biology from Bar Ilan University.

    masci2008@gmail.com

    Karmi Oxman

    Laboratory Technician

    Karmi has been a technician in the lab since June 2013. She is interested in nutrient balancing and foraging preferences of honey bees, as well as the transfer of information via honey bee dances. She has a B.A. in Biology and Environmental Studies from Oberlin College. Karmi is also an illustrator .

  • Karmi's illustrations and CV
  • LinkedIn
  • karmi.oxman@gmail.com

    Undergraduate Students


    Daniel Menchem


    Dan Eliahu


    Dror Friedman

    Lab Alumni


    Shlomi Zarchin

    Co-supervisor: Dr. Arnon Dag

    Shlomi's M.Sc. thesis was on the ability of honey bee (Apis mellifera) foragers to discriminate between pollens according to their fatty acid composition

    shlomi.zarchin@mail.huji.ac.il

    Nadav Ezra

    Nadav's M.Sc. thesis tested the subjective evaluation by honey bees of nectar secondary compounds and their fitness consequences. Specifically, he tested the effect on colony fitness of various components of avocado nectar, which is known to be unattractive to bees. In addition, he assessed the subjective evaluation of honey bees of caffeine in nectar, through analyses of their round dances.

    ndvezra@gmail.com

    Dr. Dorit Avni

    Co-supervisor: Dr. Arnon Dag

    PhD Thesis: The effect of pollen protein content and fatty acid composition on honey bee (Apis mellifera) nutrition

    avni1dorit@gmail.com

    Dr. Gal Sapir

    Co-supervisor: Prof. Martin Goldway

    PhD Thesis: Gametophytic self incompatibility in Japanese plum (Prunus salicina) – Investigation of the system and its horticultural impact

    gals@migal.org.il

    Anat Haya Zisovich

    M.Sc. Thesis: Study of pollination and fertilization factors influencing yield in the Israeli pear orchard


    Dr. Mor Salomon-Botner

    Former Postdoctoral Fellow:

    Honey bee discrimination between omega-3 and omega-6 volatiles of fatty acids

    mor.slmn@gmail.com

    Dr. Roy Kaspi

    Former Postdoctoral Fellow:

    Associative olfactory learning of the red dwarf honey bee, Apis florea

    roy.kaspi@gmail.com

    Nitzan Paldi

    M.Sc. Thesis: Using “scent genes” to enhance honey bee-assisted cross-pollination between cross-fertilizing cultivars, to produce hybrid seed


    Alon Bilu

    M.Sc. Thesis: Use of honey bees to disseminate Trichodex (Trichoderma harziatum T39) to strawberry for the control of gray mold (Botrytis cinerea)


    Yeshurun Plesser

    M.Sc. Thesis: Factors that affect the attraction of bumblebees (Bombus terrestris) to the cotton plant (Gossypium spp.)

    yplesser@biobee.com

    Lia Vaknin-Yehonatan

    M.Sc. Thesis: Comparative evaluations of reward dimensions in honey bees: evidence from two-alternative forced choice proboscis-extension conditioning

    liaguy@hotmail.com

    Sharon Golan

    M.Sc. Thesis: Risk-sensitivity in delay to reward in the honey bee (Apis mellifera)


    Tamar Drezner-Levi

    M.Sc. Thesis: Evaluation of reward distribution by honey bee (Apis mellifera) foragers collecting nectar, pollen and water

    tamarlevy2@gmail.com

    Gil Menda

    M.Sc. Thesis: An experimental study of choice behavior of honey bees in response to variability in reward magnitude using a proboscis extension response (PER) paradigm.

    gm234@cornell.edu

    Dr. Ohad Afik

    M.Sc. Thesis: The effect of ambient temperature on the crop loading behavior of honey bees (Apis mellifera)

    PhD Thesis: Factors influcencing the attraction of honey bees (Apis mellifera) to avocado (Persea americana) flowers

    ohad.afik@mail.huji.ac.il

    Dr. Arnon Dag

    Former Postdoctoral Fellow:

    Honey bee (Apis mellifera) strains differ in avocado (Persea americana) nectar foraging preferences

    arnondag@volcani.agri.gov.il